1·Leaves alternate, petiolate; leaf blade ovate to linear, margin entire or nearly so.
叶互生，具叶柄；叶片卵形到线形，边缘全缘或近全缘。
2·Leaf blade with both surfaces scaly, scales entire, or lobed.
叶片具两面有鳞的，全缘的鳞片，或浅裂的。
3·Leaf blade and petioles puberulent to woolly at least along veins; anthers globose or reniform.
对具绵状毛的至少沿脉被微柔毛的叶片和叶柄；花药球状的或肾形。
4·Leaf blade veins forming square tessellation.
叶片脉形成广场棋盘花纹。
5·Leaf blade base broadly cuneate to rounded, margin obtusely serrate; drupe and endocarp ellipsoid to oblong.
叶片基部宽楔形到圆形，边缘钝锯齿；核果和内果皮椭圆形的到长圆形。